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Nepenthes alisaputrana : ウィキペディア英語版
Nepenthes × alisaputrana

''Nepenthes × alisaputrana'' ( preferably, or , after Datuk Lamri Ali), or the Leopard Pitcher-Plant,〔Phillipps, A. & A. Lamb 1996. ''Pitcher-Plants of Borneo''. Natural History Publications (Borneo), Kota Kinabalu.〕 is a hybrid of two well-known ''Nepenthes'' pitcher plant species: ''N. burbidgeae'' and ''N. rajah''. The plant is confined to Mount Kinabalu in Sabah, Borneo.〔Clarke, C.M. 1997. ''Nepenthes of Borneo''. Natural History Publications (Borneo), Kota Kinabalu.〕
''Nepenthes × alisaputrana'' was described in 1992 by J. H. Adam and C. C. Wilcock and is named in honour of Datuk Lamri Ali, a former Director of Sabah Parks.〔 It is only known from a few remote localities within Kinabalu National Park, where it grows in stunted, open vegetation over serpentine soils at around 2000 m above sea level, often amongst populations of ''N. burbidgeae''.〔Clarke, C.M. 2001. ''A Guide to the Pitcher Plants of Sabah''. Natural History Publications (Borneo), Kota Kinabalu.〕 It grows alongside both parent species on Pig Hill,〔Thong, J. 2006. ''Victorian Carnivorous Plant Society Journal'' 82: 6–12.〕 where it is found at 1930–1950 m.〔Adam, J.H., C.C. Wilcock & M.D. Swaine 1992. ''Journal of Tropical Forest Science'' 5(1): 13–25.〕
This plant is notable for combining the best characters of both parent species, not least the size of its pitchers, which rival those of ''N. rajah'' in volume (≤35 cm high, ≤20 cm wide).〔 The other hybrids involving ''N. rajah'' do not exhibit such impressive proportions. The pitchers of ''N. × alisaputrana'' can be distinguished from those of ''N. burbidgeae'' by a broader peristome, larger lid and simply by their sheer size. The hybrid differs from its other parent, ''N. rajah'', by its lid structure, indumentum of short, brown hairs, narrower and more cylindrical peristome, and pitcher colour, which is usually yellow-green with red or brown flecking. For this reason, Anthea Phillipps and Anthony Lamb gave it the common name "Leopard Pitcher-Plant".〔 The peristome is green to dark red and striped with purple bands. Leaves are often slightly peltate. The hybrid is a strong climber and frequently produces upper pitchers.〔
''Nepenthes × alisaputrana'' more closely resembles ''N. rajah'' than ''N. burbidgeae'', but it is difficult to confuse this plant with either. However, this mistake has previously been made on at least one occasion; a pitcher illustrated in Adrian Slack's ''Insect-Eating Plants and How to Grow Them'' as being ''N. rajah''〔Slack, A. 1986. ''Insect-Eating Plants and How to Grow Them''. Alphabooks, Dorset, UK.〕 is in fact ''N. burbidgeae × N. rajah''.〔
In 2002, phytochemical screening and analytical chromatography were used to study the presence of phenolic compounds and leucoanthocyanins in ''N. × alisaputrana'' and its putative parent species.〔Adam, J. H., R. Omar & C. C. Wilcock 2002. ''OnLine Journal of Biological Sciences'' 2(9): 623–625. 〕 The research was based on leaf material from nine dry herbarium specimens. Eight spots containing phenolic acids, flavonols, flavones, leucoanthocyanins and 'unknown flavonoid' 1 and 3 were identified from chromatographic profiles. The distributions of these in the hybrid ''N. × alisaputrana'' and its putative parental species ''N. burbidgeae'' and ''N. rajah'' are shown in the table to the left. A specimen of ''N. × alisaputrana'' grown from tissue culture (''in vitro'') was also tested.〔
Luteolin, cyanidin and 'Unknown Flavonoid 3' were undetected in ''N. burbidgeae'', while concentrations of 'Unknown Flavonoid 1' were found to be weak. Chromatographic patterns of the ''N. × alisaputrana'' samples studied showed complementation of its putative parental species.〔
Myricetin was found to be absent from all studied taxa. This agrees with the findings of previous authors〔Jay, M. & P. Lebreton 1972. Chemotaxonomic research on vascular plants. The flavonoids of Sarraceniaceae, Nepenthaceae, Droseraceae and Cephlotaceae, a critical study of the order Sarraceniales. ''Naturaliste Canadien'' 99: 607–613.〕〔Som, R.M. 1988. Systematic studies on ''Nepenthes'' species and hybrids in the Malay Peninsula. Ph.D. thesis, Fakulti Sains Hayat, Universiti Kebangsaan Malaysia, UKM Bangi, Selangor Darul Ehsan.〕 and suggests that the absence of a widely distributed compound like myricetin among the ''Nepenthes'' examined might provide additional diagnostic information for these taxa.〔
==References==


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